Zbozinek, T. D., Perez, O. D., Wise, T., Fanselow, M., & Mobbs, D. (in revision, PLOS Computational Biology). Ambiguity Drives Higher-Order Pavlovian Learning. PsyArXiv

Tarantola, T., Folke, T., Boldt, A., Perez, O.D., & De Martino, B. (in revision, PLOS Computational Biology). Confirmation bias optimizes reward learning. bioRxiv.



Perez, O. D. (2022). Action sequences, habits and attention in copying strategies. Behavioral and Brain Sciences.

Perez, O. D., Vogel, E.H, Narasiwodeyar, S., & Soto, F.A. (2022). Sub-sampling of cues in associative learning. Learning & Memory.

Pool, E. R.*, Gera, R.*, Fransen, A.*, Perez, O. D.*, Cremer, A.*, Aleksic, M.*, Tanwisuth, S.*, Quail, S.*, Ceceli, A. O., Manfredi, D. A., Nave, G, Tricomi, E., Balleine, B., Schonberg, T., Schwabe, L., O'Doherty, J. P. (2022). Determining the effects of training duration on the behavioral expression of habitual control in humans: a multi-laboratory investigation. Learning & Memory.

(* Equal contribution)


Zbozinek, T. D., Wise, T., Qi, S., Perez, O. D., Fanselow, M., & Mobbs, D. (2021). Occasion Setting, Anxiety, and Their Computation in Human Fear Conditioning. Behaviour Research & Therapy.

Perez, O. D. (2021) Instrumental behavior in humans is sensitive to the correlation between response rate and reward rate. Psychonomic Bulletin and Review, 28, 649–656.


Perez, O. D. & Dickinson, A. (2020). A theory of actions and habits: The interaction of rate correlation and contiguity systems in free-operant behavior. Psychological Review, 127 (6), 945–971.

Perez, O. D. & Soto, F.A. (2020). Evidence for a dissociation between causal beliefs and instrumental actions. Quarterly Journal of Experimental Psychology, 73 (4), 495-503. https://doi:10.31234/


Dickinson, A. & Perez, O. D. (2019). Actions and Habits: Psychological issues in Dual-System Theory. In R.W. Morris & A. Bornstein (Eds.) Goal-Directed Decision Making: Computations and Neural Circuits. Elsevier. doi:10.1016/B978-0-12-812098-9.00001-2

Perez, O. D., Aitken, M. R., Zhukovsky, P., Soto, F. A., Urcelay, G. P., & Dickinson, A. (2019). Human instrumental performance in ratio and interval contingencies: A challenge for associative theory. Quarterly Journal of Experimental Psychology, 72(2), 311-321.


Pauli, W. M., Cockburn, J., Pool, E. R., Perez, O. D., & O’Doherty, J. P. (2018). Computational approaches to habits in a model-free world. Current Opinion in Behavioral Sciences, 20, 104-109.

Perez, O. D., San Martin, R., & Soto, F. A. (2018). Exploring the Effect of Stimulus Similarity on the Summation Effect in Causal Learning. Experimental Psychology, 65, 183-200. doi:10.1027/1618-3169/a000406

Perez, O. D., Aitken, M. R. F., Milton, A. L., & Dickinson, A. (2018). A re-examination of responding on ratio and regulated-probability interval schedules. Learning and Motivation, 64, 1-8.

In preparation

Perez, O. D., & Dickinson, A. Dual-system avoidance.

Perez, O. D., Oh, S., Dickinson, A., & Merlo, E. The effect of extinction and causal degradation on the goal-directed status of an action.